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Analyses of Phylogenetics and Evolution

300packages dependsuggested by 100
1.5Mdownloads / year
–test coverage
13/13flavors OK

Overview

About

Functions for reading, writing, plotting, and manipulating phylogenetic trees, analyses of comparative data in a phylogenetic framework, ancestral character analyses, analyses of diversification and macroevolution, computing distances from DNA sequences, reading and writing nucleotide sequences as well as importing from BioConductor, and several tools such as Mantel's test, generalized skyline plots, graphical exploration of phylogenetic data (alex, trex, kronoviz), estimation of absolute evolutionary rates and clock-like trees using mean path lengths and penalized likelihood, dating trees with non-contemporaneous sequences, translating DNA into AA sequences, and assessing sequence alignments. Phylogeny estimation can be done with the NJ, BIONJ, ME, MVR, SDM, and triangle methods, and several methods handling incomplete distance matrices (NJ*, BIONJ*, MVR*, and the corresponding triangle method). Some functions call external applications (PhyML, Clustal, T-Coffee, Muscle) whose results are returned into R.

Install

Health

CRAN checks
13OK
Slowest check: 7.8 min · r-oldrel-windows-x86_64
Status changes
  • OK13 OK2026-09-08
  • ERROR12 OK, 1 ERROR2026-09-07
  • OK13 OK2026-08-05
  • NOTE12 OK, 1 NOTE2026-08-01
  • OK12 OK2026-04-25
Show 3 earlier changesHide 3 earlier changes
  • NOTE12 OK, 2 NOTE2026-04-22
  • ERROR11 OK, 2 NOTE, 1 ERROR2026-04-18
  • NOTE12 OK, 2 NOTE2026-03-10
Advisories

none known

Documentation

In release 5.8-1
READMEYes · 15 wordsVignettesYes · 3 · dynamicNEWSYes · 0% structuredCitation fileYes · cites a publicationWebsitenot declared in DESCRIPTION
Help pages
188 (1 internal)includes a package overview page
Examples that run
90%131 run in a plain check · 13 never run · 1 hold no code
Exported functions with examples
80%
Documented parameters
94%
Return-value docs
91%
Help pages citing references
51%

Downloads

1.5M
CRAN downloads in the past year
Rank #307 · ~4K/day · ~123.1K/mo
Daily download trend is not available in this view yet.
126K30 days
358.9K90 days
1.5M1 year
Compare downloads with other packages →
Also on26.3K r2u187 autocran705.6K conda_forge1.2K c2d4u

Repository

75Stars
14Forks
7Open issues
0Open PRs
0Releases
284Commits
12Commit identities
License file reads as GPL-2.0 · 284 commits · Last activity 2026-10-20 · +4.2% stars, 30d · Default branch at 5.8-3, ahead of CRAN 5.8-1 (checked 2026-10-18)
Responsiveness
How these are sampled
PRs merged (all time)
64%
Median days to close, 50 most recently updated closed issues
10
Median days to close, 50 most recently updated closed PRs
6

Stars over time

2025-09-21 · 722026-10-15 · 75
PracticesBeta

2 development-tooling and community-health practices detected across 1 family in the upstream repository

Checks run against github.com/emmanuelparadis/ape on 2026-10-18.

Continuous integration (2)
GitHub ActionsR CMD check workflow

The R CMD check workflow runs on Linux, macOS and Windows, and on R-devel

How this is detected·Detection ruleset v3 (2026-09-26)

Dependencies

Declared dependencies
8 external dependencies (excludes base and recommended)

Installing ape adds 2 packages (2 with compiled code, 2.0 MB of source tarballs). Heaviest: Rcpp 1.8 MB, digest 231 KB.

Depends (1)
R >= 3.2.0
Imports (9)
nlmelatticegraphicsmethodsstatsutilsparallelRcppdigest
LinkingTo (1)
Enhances (0)
none

Datasets

People & History

People (33)
Author, Maintainer, Copyright holder · added in 1.2-7
Author, Copyright holder · added in 3.2
Author, Copyright holder · added in 1.2-7
Author, Copyright holder · added in 5.0
Author, Copyright holder · added in 5.4
Author, Copyright holder · added in 1.2-7
Author, Copyright holder · added in 2.0
Author, Copyright holder · added in 2.0
Author, Copyright holder · added in 3.2
Author, Copyright holder · added in 1.10-1
Author, Copyright holder · added in 1.2-7
Author, Copyright holder · added in 4.1
Author, Copyright holder · added in 2.0
Author, Copyright holder · added in 5.1
Author, Copyright holder · added in 2.0
Author, Copyright holder · added in 3.3
Author, Copyright holder · added in 4.0
Author, Copyright holder · added in 5.0
Author, Copyright holder · added in 2.3
Author, Copyright holder · added in 2.0
Author, Copyright holder · added in 2.3
Author, Copyright holder · added in 1.8-3
Author, Copyright holder · added in 5.4
Author, Copyright holder · added in 5.2
Author, Copyright holder · added in 5.8-1
Author, Copyright holder · added in 3.1-3
Author, Copyright holder · added in 1.10-1
Author, Copyright holder · added in 1.2-7
Author, Copyright holder · added in 2.8
Author, Copyright holder · added in 3.2
Author, Copyright holder · added in 2.6
Author, Copyright holder · added in 1.2-7
Author, Copyright holder · added in 2.2

Maintainer since 0.1 (2002)

Listed in earlier versions (8)
Show 8 peopleHide 8 people
Emmanuel Paradis Julien Claude Yvonnick Noel Ben Bolker
no longer listed · 0.1 to 0.2
Emmanuel Paradis Korbibian Strimmer Julien Claude Yvonnick Noel Ben Bolker
no longer listed · 0.2-1
Emmanuel Paradis Korbinian Strimmer Julien Claude Yvonnick Noel Ben Bolker
no longer listed · 1.0
Emmanuel Paradis Korbinian Strimmer Julien Claude Gangolf Jobb Yvonnick Noel Ben Bolker
no longer listed · 1.1 to 1.2-3
Emmanuel Paradis Korbinian Strimmer Julien Claude Gangolf Jobb Rainer Opgen-Rhein Yvonnick Noel Ben Bolker
no longer listed · 1.2-4 to 1.2-5
Emmanuel Paradis Korbinian Strimmer Julien Claude Gangolf Jobb Rainer Opgen-Rhein Julien Dutheil Yvonnick Noel Ben Bolker
no longer listed · 1.2-6
Gangolf Jobb
no longer listed · 1.2-7 to 2.5-1
Yvonnick Noel
no longer listed · 1.2-7 to 3.0-9
Releases (104)

Pick two releases to compare their code metrics.

  • R
    R 4.6.0 released · 2026-04-24
  • R
    R 4.5.0 released · 2025-04-11
  • 5.8-1Latest
    record
    2024-12-16 · current release · diff ↗ ·
    added 1 export
    write.phyloXML
  • R
    R 4.4.0 released · 2024-04-24
  • 5.8
    record
    2024-04-11 · diff ↗
  • R
    R 4.3.0 released · 2023-04-21
  • 5.7-1
    record
    2023-03-13 · diff ↗
  • 5.7
    record
    2023-02-16 · diff ↗ ·
    added 7 exports
    drop.tip.multiPhylo, drop.tip.phylo, efastats, keep.tip.multiPhylo, keep.tip.phylo, letterconf, muscle5
  • R
    R 4.2.0 released · 2022-04-22
  • 5.6-2
    record
    2022-03-02 · diff ↗
  • 5.6-1
    record
    2022-01-07 · diff ↗
  • 5.6
    record
    2021-12-21 · diff ↗ ·
    added 2 exports
    degree, solveAmbiguousBases
  • R
    R 4.1.0 released · 2021-05-18
  • 5.5
    record
    2021-04-25 · diff ↗ ·
    added 4 exports
    LargeNumber, SHORTwise, rmtopology, rtopology
  • 5.4-1
    record
    2020-08-13 · diff ↗ ·
    added 1 export
    nexus2DNAbin
  • 5.4
    record
    2020-06-03 · diff ↗ ·
    removed 2 exports
    neworder_phylo, neworder_pruningwise
    added 2 exports
    getAnnotationsGenBank, latag2n
Show 123 earlier eventsHide 123 earlier events

Package metadata

First published
2002-08-31
Total releases
104 / 24 yrs
License
GPL-2 | GPL-3 OSI
Minimum R
≥ 3.2.0
Bundled data
125 KB / 16 files
Download size
1.4 MB

Cite

Cite this package

Run in R for the authors' entry:

citation("ape")

The package provides a citation file, so citation() returns the authors' own entry.

It points to 10.1093/bioinformatics/bty633.

Cite the R Observatory

For a number measured here: a download total, a coverage figure, an archival date.

APA

Balamuta, J. J. (2026). R Observatory: Metrics for ape version 5.8-1 [Data set]. HJJB, LLC. Data release v2026-10-21. https://doi.org/10.5281/zenodo.21843040

From data release v2026-10-21, which the citation names so these numbers can be found later. More on citing and the projects behind them.

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